If a model doesn't even closely predict an observation, we have a problem.
Yes, the model might be inadequate.
Or, there is a fundamental flaw, or an extraordinarily significant omission, in our current understanding of evolutionary processes.
I suspect the former.
To be honest, I'm not aware of a mathematical "model of evolution" from a credible source which has caused the researchers to conclude that there must be an additional organising factor that has thus far been overlooked.
I think such a model would be extraordinarily ambitious (and highly contentious, considering our current understanding of abiogenesis and the emergence of eukaryotes, multicellular organisms and sexual reproduction).
However, I won't pretend to have done a literature trawl in this area!
I feel (LilWabbit) that you're claiming that there has been insufficient time for the current level of biological complexity to evolve according to mainstream theories- a claim which most suitably qualified people (biologists, geneticists, palaeontologists etc.) might disagree with. -That latter supposition of mine doesn't necessarily make you wrong, of course.
Your solution, to this problem that may well not exist, is to re-interpret what most people "in the field" might consider to be observed reality- essentially Darwinian gradualism, with an increased understanding of the role of small, isolated genepools in promoting speciation- to include a discrete (as opposed to emergent) driver for complexity, or at least a favouring of complexity in itself (as opposed to a favouring of traits resulting from genetic mutation or reproductive "shuffling", sometimes with concomitant genomic complexity).
Apologies if I have mis-represented your position!
Yet there's only a finite stretch of punctuated evolutionary periods during which these mutations are thought to have occurred at a certain averaged frequency to cause more complex species. Therefore the probability for even a single organism to produce mutated offspring falling within a narrow range of more complex favourable mutations is zero, let alone for their offspring to continue mutating favourably within mutant subpopulations into more complex and therefore less probable configurations.
Yet there's only a finite stretch of punctuated evolutionary periods
Even Eldredge and Gould didn't claim that all evolution occurred during "punctuation", not by a long way.
There are lots of examples of phyletic gradualism.
While there is a tendency for us to focus on the most dramatic "punctuative" events- the Cambrian explosion, the post K-T speciation- many (most) species may have undergone significant change (or indeed have evolved into a new species) as a chronospecies, without any "punctuative" events at all, or during a punctuation that primarily effects that species only and, while perhaps visible in the fossil record, might not coincide with major changes in the geological record (as evidenced in geological strata etc.)
A chance separation of a small breeding population- or even one breeding pair- from the rest of their genepool might be sufficient to enable speciation.
...which these mutations are thought to have occurred at a certain averaged frequency to cause more complex species.
By whom? And what is their evidence?
Plus, the existing variations of genotype within a species can permit evolution if selection pressures change, without mutation.
the probability for even a single organism to produce mutated offspring falling within a narrow range of more complex favourable mutations is zero
COVID-19 seemed to do a pretty good job of spreading itself about, mutations causing greater transmissibility.
Methicillin-resistant Staphylococcus Aureus has been favoured by the selection pressure of (inefficient) antibiotic use, and has spread globally.
Its even nastier offspring, vancomycin-resistant S.A., is only known from 1996- presumably driven by the use of vancomycin to treat MRSA.
Lactase persistence has evolved- probably several times in different locations, with differing mechanisms- in the past few thousand years in humans; north Europeans with the necessary mutation(s) gained a significant reproductive advantage over those who did not. The genetic basis of lactase persistence is
reasonably well understood;
Genetic Signatures of Strong Recent Positive Selection at the Lactase Gene, Bersaglieri, T., Sabeti, P.C. et al, 2004,
American Journal of Human Genetics 74 (6), link:
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1182075/
let alone for their offspring to continue mutating favourably within mutant subpopulations into more complex and therefore less probable configurations.
No! A gene may be mutated, a population is a population.
A mutation that prevents a creature from reproducing is not beneficial pretty much by definition, so a beneficial mutation must enable its bearer to reproduce. It's then down to the genetic lottery (if the creature reproduces sexually) if that mutation is passed on to its offspring, giving them a genotype which includes the parent's mutation, and a physiology (and possibly phenotype) that reflects this, i.e. it inherits the beneficial trait.
A beneficial, heritable trait will spread, all other things being equal. But creatures bearing the new beneficial trait are not somehow immune to ongoing selection pressures or spontaneous genetic mutation; their increased complexity doesn't make further changes to the genome less probable!
Your (LilWabbit's) claim (more complex = less probable) reminds me of Fred Hoyle's "Junkyard Tornado"; he thought terrestrial abiogenesis was as likely as a tornado sweeping through a junkyard and leaving behind a Boeing 747.
External Quote:
Hoyle's fallacy contradicts many well-established and widely tested principles in the field of
evolutionary biology.
[5] As the fallacy argues, the odds of the sudden construction of higher lifeforms are indeed improbable. However, what the junkyard tornado postulation fails to take into account is the vast amount of support that evolution proceeds in many smaller stages, each driven by
natural selection[6] rather than by random chance, over a long period of time.
https://en.wikipedia.org/wiki/Junkyard_tornado
Evolution is cumulative. Think of the example of eyes, which has to be repeated
ad nauseum for creationists. (Who won't listen).
If you think it's impossible for increasing complexity to arise through natural selection, you don't believe in evolution
(not the scientific theory of evolution anyway).
Article: To study the evolution of complexity in organisms, we need an understanding of complexity that enables us to measure it. In biology today, organismal complexity has two main operational senses: (1) a horizontal sense: the number of different part types at a given hierarchical level (e.g., the number of cell types in a multicellular individual) and (2) a vertical sense: the number of levels of nestedness of parts within wholes (e.g., a eukaryotic multicellular individual is one level of nestedness above a free-living protist). How do horizontal and vertical complexity behave in evolution? For horizontal complexity, an increasing trend is predicted by current theory, that is, by the zero-force evolutionary law (ZFEL), but at most hierarchical levels, evidence is lacking and the existence of a trend is uncertain.
For vertical complexity, there is unambiguous evidence for a trend in the maximum, a rise in the maximum hierarchical level achieved by organisms over the history of life. However, the underlying mechanism of change and the forces driving the trend are unknown. Interestingly, there is some evidence that the rise in vertical complexity, the addition of new levels, is – when it occurs – accompanied by systematic losses in horizontal complexity at lower levels. Source:
https://link.springer.com/referenceworkentry/10.1007/978-3-319-32979-6_123
This is from Daniel W. McShea's "Evolution of Complexity", which was published as one chapter (of 86) in
Evolutionary Developmental Biology, eds. L.N. de la Rosa and G.B. Muller, Springer Cham 2020,
link
https://link.springer.com/referencework/10.1007/978-3-319-33038-9?page=2#about-authors
His paper has been cited twice.
He states,
External Quote:
...an increasing trend [in complexity] is predicted by current theory, that is, by the zero-force evolutionary law (ZFEL)
"Zero-force evolutionary law" sounded a bit dramatic to me, and I'd never heard of it, so I had a quick surf...
...the founder of this law is the same Daniel McShea, along with Robert N. Brandon, who described it in the book
"
Biology's First Law", sub-title "
The Tendency for Diversity and Complexity to Increase in Evolutionary Systems", 2010;
you can buy it here
https://bibliovault.org/BV.landing.epl?ISBN=9780226562261
Now, I'm sure Dr McShea is a thoroughly decent chap, but I'm not sure he's quite the same rank as Darwin, Crick/ Franklin/ Watson or Pasteur.
The idea that "ZFEL" is in any sense "
Biology's First Law" (even though he does say so himself!) is, of course, ridiculous.
In fairness, McShea and Brandon didn't claim (as far as I could see) that there is a need to invoke any extra-biological agency;
they
do believe that
External Quote:
... in the absence of natural selection, an evolutionary system with
variation and
heredity will tend spontaneously to diversify and complexify.
"Zero -Force Evolutionary Law", Wikipedia,
https://en.wikipedia.org/wiki/Zero-Force_Evolutionary_Law
Quite how you can have a real-world environment without natural selection, when you have variation and heredity as axiomatic, is not explained.
McShea seems to like "complexity" although his views about it seem to have changed rather a lot; in 1996's
Perspective Metazoan Complexity and Evolution: Is There a Trend? in Evolution 1996, 50 (2), he writes
External Quote:
In metazoans as a whole, there is good evidence only for an early-Phanerozoic trend, and only in one type of complexity.
(abstract here,
https://pubmed.ncbi.nlm.nih.gov/28568940/)
No good evidence for a trend toward complexity since the early Phanerozoic?
There clearly
is a trend; what we're discussing is whether it's the result of accumulated change due to the theory of evolution,
or whether it's due to some unidentified drive.
The probability of life arising on other planets is highly debatable (and interesting) but I'm not sure invoking metaverses or whatever in discussions of life on Earth will help much.
Let's get back to "Are all UFO reports wrong..."